Meghana Raj first debuted as a theater artist, when she was a child, acting alongside her father Sunder Raj in the play Jokumaraswamy. During the audio launch function of the Tamil film Poi in February 2006, she was invited by its director K. Balachander to pursue an acting career, starring in one of his forthcoming films. Accepting his offer, she was given the lead female role in the Tamil film Krishnaleelai, a Balachander production, directed by Selvan. However, despite continuing shooting throughout 2008, which was completed in early 2009, the film got stuck in production hell, and its release got delayed for various reasons.
Even before the release of that film, she commenced shooting for a Telugu film Bendu Apparao R.M.P, which would eventually be her first feature film release. She played the character of a school teacher in the comedy entertainer, starring Allari Naresh and Kamna Jethmalani as well, which released in October 2009 and went on to become highly successful at the box office. She debuted in the Kannada films by enacting the female lead role in Punda, a remake of the 2007 Tamil film Polladhavan.Meghana's next release was her debut Tamil film, the Boopathy Pandian-directed Kaadhal Solla Vandhen, which was a commercial failure, while she also signed up her first Malayalam project, the horror film Yakshiyum Njanum under Vinayan's direction, which was released in August 2010.The film was an average grosser, but its glamorously picturized songs on Meghana was a hit among the youth. During that time she also acted in two more Tamil projects, Nandha Nanditha, which is a remake of the 2008 Kannada film Nanda Loves Nanditha and Kalla Siripazhaga.
After a series of flops for most of 2011 where she was seen repeatedly cast merely as a glamour actress, Meghana reverted to winning ways with the Malayalam film Beautiful which was released in the first week of December. The film was a success both commercially and critically and critics widely regarded it as one of the best Malayalam movies of 2011. Meghana's portrayal of Anjali, a home nurse in the film won her very good reviews. Since getting noticed with Beautiful, Meghana got a lot of Malayalam films offers.
Meghana then underwent a total physique revamp by going through a strict diet and exercise regime. She made this decision partly because she was getting constantly ignored in Kannada films (her mother tongue) and the reason told was she was fat.
Her 2012 Malayalam film Namukku Parkkan did a decent business at the box office. Subsequent films Mullamottum Munthiricharum, Banking Hours 10 to 4, Maad Dad, 'Madirashi and 'Red Wine failed to perform. However, her next film Memories opposite Prithviraj Sukumaran opened to overwhelmingly positive reviews and was a success.
Meghana Raj repeated the success in her next release Raja Huli in Kannada. The film opened to rave reviews and has been declared a super hit.
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